1. Department of Marine Biological Science and Technology, College of Ocean and Earth Sciences, Xiamen University, Xiamen, Fujian, China
2. Marine and Fishery Institute of Xiamen, Xiamen, Fujian, China
3. School of Biology and Ecology, Department of Mathematics and Statistics, University of Maine, Orono, Maine, USA
4. College of Science and Engineering, Centre for Sustainable Tropical Fisheries and Aquaculture, James Cook University, Townsville Australia
5. Fisheries College, Jimei University, Xiamen, Fujian, China