1. Molecular Invertebrate Systematics and Ecology Lab, Graduate School of Engineering and Science, University of the Ryukyus, Nishihara, Nishihara, Japan
2. Graduate School of Engineering and Science, University of the Ryukyus, Nishihara, Okinawa, Japan
3. Tropical Biosphere Research Center, University of the Ryukyus, Nishihara, Okinawa, Japan