1. Jiangsu Key Laboratory of Marine Bioresources and Environment, Jiangsu Ocean University, Lianyungang, China
2. Co-Innovation Center of Jiangsu Marine Bio-industry Technology, Jiangsu Ocean University, Lianyungang, China
3. Scripps Institution of Oceanography, University of California San Diego, La Jolla, CA, USA
4. Jiangsu Marine Resources Development Research Institute, Jiangsu Ocean University, Lianyungang, China