1. Department of Ophthalmology, University of Pittsburgh School of Medicine, Pittsburgh, PA, USA;
2. Department of Ophthalmology, Wilmer Eye Institute, the Johns Hopkins University School of Medicine, Baltimore, MD, USA
3. Department of Cell Biology and Center for Biologic Imaging, University of Pittsburgh School of Medicine, Pittsburgh, PA, USA;
4. Kodiak Sciences, Palo Alto, CA, USA;
5. Institut De La Vision, INSERM, CNRS, Sorbonne Université, Paris, France