1. Department of Archaeology and Anthropology, Division of Biological Anthropology, University of Cambridge, Cambridge, UK;
2. McDonald Institute of Archaeological Research, University of Cambridge, Cambridge, UK;
3. Department of Genetics, Centre for Microbial Ecology and Genomics, Genomic Research Institute, University of Pretoria, Hatfield, South Africa;
4. Department of Anthropology & Geography, Human Origins and Palaeoenvironmental Research Group (HOPE), Oxford Brookes University, Oxford, UK;
5. Leverhulme Centre for Human Evolutionary Studies, Henry Wellcome Building, Cambridge, UK