1. Ottawa Research and Development Centre, Agriculture and Agri-Food Canada, 960 Carling Ave. Ottawa, ON K1A 0C6, Canada
2. Department of Botany Faculty of Science, Charles University, Benatska 2CZ-128 01 PRAHA 2, Prague, Czech Republic
3. Department of Environmental Science, Ishikawa Prefectural University, 1–308 Suematsu, Nonoichi, Ishikawa 921-8836, Japan