1. Faculty of Applied Biological Sciences, Gifu University, Yanagido 1-1, Gifu City, Gifu 501-1193, Japan
2. Department of Biology, College of Science, Taif University, P.O. Box 11099, Taif, 21944, Saudi Arabia
3. Department of Genetics, Faculty of Agriculture, Menoufia University, Sheben El-Kom 51132, Egypt
4. Agricultural Botany Department, Faculty of Agriculture, Kafrelsheikh University, Kafr Elsheikh, 33516, Egypt