1. Higher Institute of Agriculture of Chott-Mariem, University of Sousse, Sousse 4042, Tunisia
2. Laboratory of Management and Valorization of Forest Resources, National Research Institute of Rural Engineering, Water and Forests, University of Carthage, Ariana 2080, Tunisia
3. National Institute of Applied Sciences and Technology (INSAT), University of Carthage , Tunis Cedex BP 676 - 1080, Tunisia