1. Dorset Environmental Science Centre, Ontario Ministry of the Environment, Conservation and Parks, Dorset, ON, P0A 1E0, Canada;
2. Associated Environmental Consultants Inc, Edmonton, AB, T5J 5C6, Canada;
3. Natural Resources Research Institute, University of Minnesota Duluth, Duluth, MN 55811, USA;
4. Department of Integrative Biology, University of South Florida, St. Petersburg, FL 33701, USA