1. a Department of Veterinary Medicine , College of Bioresource Sciences, Nihon University , Fujisawa , Kanagawa , 252‐8510 , Japan
2. b Laboratory of Developmental and Reproductive Biotechnology, Department of Animal Science and Biotechnology, College of Veterinary Medicine, Azabu University, 1-17-71, Fuchinobe , Sagamihara , Kanagawa , 229‐8501 , Japan
3. c Faculty of Agriculture , University Putra Malaysia , Serdang , Selangor , 43400 , Malaysia
4. d Center for Ecological Research , Kyoto University, Hiranocho , Otsu , Shiga , 520‐2113 , Japan
5. e Department of Zoology , National Science Museum, Tokyo , Shinjuku , Tokyo , 169‐0073 , Japan
6. f Laboratory of Animal Morphology and Function, Graduate School of Bioagricultural Sciences, Nagoya University, Furocho , Chikusa , Nagoya , 464‐8601 , Japan