1. a Triticeae Research Institute, Sichuan Agricultural University, Wenjiang , Chengdu , 611130 , Sichuan , China
2. b Key Laboratory of Crop Genetic Resources and Improvement, Ministry of Education , Sichuan Agricultural University , Yaan , 625014 , Sichuan , China
3. d Biology and Microbiology Department , South Dakota State University , Brookings , SD 57007 , USA
4. c Northwest Plateau Institute of Biology, Chinese Academy of Science , Qinghai , 810001 , China