1. a Laboratory of Molecular Ecology, Department of Biology, Tunghai University, Taichung , Taiwan , 407 , R. O. C.
2. b Department of Biofunctional Science, Faculty of Agriculture and Life Science, Hirosaki University , Hirosaki , 036-8561 , Japan
3. c Laboratory of Wildlife Ecology, Department of Biology,Tunghai University,Taichung , Taiwan , 407 , R. O. C
4. d Chromosome Research Unit, Faculty of Science, Hokkaido University , Sapporo , 060-0810 , Japan