1. a Graduate School of Human Health Science , Kochi Women’s University, 5-15 Eikokuji-cho , Kochi , 780-8515 , Japan
2. b Herbarium (HAST), Biodiversity Research Center , Academia Sinica , Taipei 115 , Taiwan
3. c Department of Plant Science, School of Agriculture , Tokai University, Kawayou, Minamiaso-mura, Aso-gun , Kumamoto , 869-1404 , Japan
4. d Graduate School of Human and Environmental Studies , Kyoto University, Yoshidanihonmatsu-cho, Sakyo-ku , Kyoto , 606-8501 , Japan
5. e Laboratory of Ecology and Systematics, Faculty of Science , University of Ryukyus , 1 Senbaru, Nishihara-cho, Nakagami-gun, Okinawa , 903-0213 , Japan
6. f Department of Biology, Faculty of Human Life and Environmental Science , Kochi Women’s University, 5-15 Eikokuji-cho , Kochi , 780-8515 , Japan