1. a College of Pharmacy , Wuhan University , Wuhan , 430072 , China
2. d College of Life Science and Technology , Chongqing University of Arts and Sciences , Chongqing , 402168 , China
3. b Institute of Plant Genetics and Breeding, School of Biological Technology and Engineering, Guizhou Normal University , Guiyang , 550001 , China
4. e Institute of Chinese Herbal Medicines, Hubei Academy of Agricultural Sciences , Enshi , 445000 , China
5. c Key Lab of the Ministry of Education for Plant Developmental Biology , College of Life Science, Wuhan University , Wuhan , 430072 , China