1. The United Graduate School of Agricultural Science, Gifu University, Gifu, Japan
2. Graduate School of Natural Science and Technology, Gifu University, Gifu, Japan
3. Experimental Plant Division, RIKEN BioResource Research Center, Tsukuba, Ibaraki, Japan
4. RIKEN CSRS, Suehiro-cho, Tsurumi-ku, Yokohama, Japan