1. Institute of Transformative Bio-Molecules (WPI-ITbM), Nagoya University, Furo-cho, Chikusa-ku, Nagoya, Japan
2. Graduate School of Biological Sciences, Nara Institute of Science and Technology, Takayama, Ikoma, Japan
3. Department of Biology, University of Washington, Seattle, WA, USA
4. Howard Hughes Medical Institute, University of Washington, Seattle, WA, USA
5. Division of Biological Science, Graduate School of Science, Nagoya University, Furo-cho, Chikusa-ku, Nagoya, Japan