1. Laboratory of Plant Molecular Genetics; Graduate School of Biological Sciences; Nara Institute of Science and Technology; Ikoma, Japan
2. Cell and Developmental Biology; John Innes Center; Norwich, United Kingdom
3. These authors equally contributed to this work.
4. Deptartment of Biological Science; University Tunku Abdul Rahman Jalan Universiti; Bandar Barat Kampar, Malaysia
5. Shanghai Center for Plant Stress Biology; Shanghai, PR China