1. a Department of Forest Soils , Swedish University of Agricultural Sciences , Box 7001, Uppsala, S‐750 07, Sweden
2. b Department of Biology , Mount Union College , Alliance, Ohio, 44601, USA
3. c Dipartimento Biologia Vegetale , Universita di Napoli , Via Foria 223, Napoli, 1–80139, Italy
4. d Department of Statistics , Swedish University of Agricultural Sciences , Box 7013, Uppsala, S‐750 07, Sweden