1. School of Life and Environmental Sciences, University of Lincoln, Lincoln, UK
2. Department of Classics and Archaeology, University of Nottingham, Nottingham, UK
3. Department of Biological Sciences, University of Cape Town, Rondebosch, South Africa
4. School of Life Sciences, University of KwaZulu-Natal, Pietermaritzburg, South Africa
5. School of Veterinary and Life Sciences, Murdoch University, Murdoch, WA, Australia