1. Guangzhou Key Laboratory of Subtropical Biodiversity and Biomonitoring, Guangdong Provincial Key Laboratory of Healthy and Safe Aquaculture, College of Life Science, South China Normal University, West 55 of Zhongshan Avenue, Guangzhou, 510631, Guangdong Province, China
2. Natural History Museum of Denmark, University of Copenhagen, Øster Farimagsgade 5, Copenhagen K 1307, Denmark
3. The Eighth Geological Brigade, Hebei Geological Prospecting Bureau, Yanshan Street 221, Qinhuangdao, 066001, Hebei Province, China