1. Aquatic Plant Variety Center, National Institute of Fisheries Science, Tongildaero 130, Mokpo, Jeollanamdo, 58746, Korea
2. Field Science Center for Northern Biosphere, Hokkaido University, Sapporo, Hokkaido 060-0809, Japan
3. Key Laboratory of Experimental Marine Biology, Institute of Oceanology, Chinese Academy of Sciences, 7 Nanhai Road, Qingdao, Shandong 266071, China
4. Center for Ocean Mega-Science, Chinese Academy of Sciences, 7 Nanhai Road, Qingdao, Shandong 266071, China
5. Laboratory for Marine Biology and Biotechnology, Qingdao National Laboratory for Marine Science and Technology, 7 Nanhai Road, Qingdao, Shandong 266071, China