1. Takehara Fisheries Research Station, Setouchi Field Science Center, Graduate School of Integrated Sciences for Life, Hiroshima University, Takehara, Hiroshima 725-0024, Japan
2. Department of Fishery Biology, Faculty of Fisheries, Kasetsart University, Bangkok 10900, Thailand
3. Graduate School of Integrated Science for Life, Hiroshima University, Higashi-Hiroshima, Hiroshima 739-8528, Japan
4. Central Laboratory, Marine Ecology Research Institute, Iwawada, Onjuku-machi, Isumi-gun, Chiba 299-5105, Japan