1. Department of Botany and Beaty Biodiversity Research Centre, University of British Columbia, 3200-6270 University Boulevard, Vancouver, British Columbia, V6T 1Z4, Canada
2. University Herbarium, University of California at Berkeley, 1001 Valley Life Sciences Building #2465, Berkeley, CA 94720-2465, USA
3. Division of Mathematics, Science, and Engineering, Hartnell College, 411 Central Ave., Salinas, California, 93901, USA
4. Biology Department and Herbarium, University of North Carolina Chapel Hill, Coker Hall CB 3280, Chapel Hill, North Carolina, 27599-3280, USA