1. Institute of Oceanography and Environmental Science, Mindanao State University, Tawi-Tawi College of Technology and Oceanography, Boheh Sallang, Sanga-Sanga, Bongao, Tawi-Tawi 7500, Philippines
2. Institute of Marine Biology, National Taiwan Ocean University, Keelung 20224, Taiwan, R.O.C.
3. School of Biological Sciences, Victoria University of Wellington, Wellington 6012, New Zealand
4. School of Biosciences, University of Melbourne, Parkville, Victoria 3010, Australia
5. Tasmanian Herbarium, College Road, Sandy Bay, Tasmania 7005, Australia