1. Department of Biodiversity and Conservation Biology, University of the Western Cape, P. Bag X17, Bellville 7535, South Africa
2. Australian Rivers Institute-Coast & Estuaries, School of Environment and Science, Nathan Campus, Griffith University, 170 Kessels Road, Nathan QLD 4111, Australia
3. Department of Life Science, Chosun University, 309 Pilmun-daero, Dong-gu, Gwangju 61452, Korea
4. Division of Mathematics, Science, and Engineering, Hartnell College, 411 Central Ave., Salinas, California 93901, USA
5. Biology Department and Herbarium, University of North Carolina at Chapel Hill, Coker Hall CB 3280, Chapel Hill, North Carolina 27599-3280, USA