1. Faculty of Marine Science and Technology, Fukui Prefectural University, 1-1 Gakuencho, Obama, Fukui, 917-0003, Japan
2. Science Division, Royal Botanic Garden, Edinburgh, 20A Inverleith Row, Edinburgh, EH3 5LR, Scotland, UK
3. Marine and Continental Waters Programme, Institute for Food and Agricultural Research and Technology (IRTA), Sant Carles de la Ràpita, Catalunya, E-43540, Spain