1. Key Laboratory of Marine Ecosystem Dynamics, Second Institute of Oceanography, Ministry of Natural Resources, Hangzhou 310012, China
2. College of Agriculture & Biotechnology, Zhejiang University, Hangzhou 310058, China
3. Institute of Marine Science Ocean College, Zhejiang University, Zhoushan 316021, China
4. Key Laboratory of Tropical Marine Ecosystem and Bioresource, Fourth Institute of Oceanography, Ministry of Natural Resources, Beihai 536007, China
5. School of Life Sciences, Xiamen University, Xiamen 361102, China