1. Division of Plant Science, University of Dundee at the James Hutton Institute, Invergowrie, Dundee DD2 5DA, UK
2. Climate Change Cluster, Faculty of Science, University of Technology, Sydney, Sydney, Ultimo 2007, Australia
3. School of Biological Sciences, University of Western Australia, Crawley 6009, Australia
4. School of Geographical Sciences, University of Bristol, Bristol BS8 1SS, UK