1. CAS and Shandong Province Key Laboratory of Experimental Marine Biology, Center for Ocean Mega-Science, Institute of Oceanology, Chinese Academy of Sciences, No. 7 Nanhai Road, Qingdao, 266071, China
2. Laboratory for Marine Biology and Biotechnology, Qingdao National Laboratory for Marine Science and Technology, No. 168 Wenhai Road, Qingdao, 266071, China
3. College of Earth and Planetary Sciences, University of Chinese Academy of Sciences, No. 19 Yuquan Road, Beijing, 100049, China