1. Marine Research Centre, Finnish Environment Institute (SYKE), Agnes Sjöbergin katu 2, Helsinki 00790, Finland
2. Marine Biological Section, University of Copenhagen, Universitetsparken 4, Copenhagen Ø 2100, Denmark
3. Department of Bioscience, Aarhus University, Frederiksborgvej 399, Roskilde 4000, Denmark
4. Natural History Museum of Denmark, University of Copenhagen, Øster Farimagsgade 5, Copenhagen K 1353, Denmark