1. Department of Botany and Plant Pathology, College of Agricultural Sciences, Oregon State University, 2082 Cordley Hall, Corvallis, Oregon 97331-2902, USA
2. Institute for East China Sea Research (ECSER), Nagasaki University, 1551-7, Taira-machi, Nagasaki 851-2213, Japan
3. LITTORAL, Ifremer, Concarneau F‐29900, France