1. College of Marine Ecology and Environment, Shanghai Ocean University, No.999, Huchenghuan Road, Pudong New District, Shanghai 201306, China
2. College of Fisheries, Guangdong Ocean University, No.1, Haida Road, Mazhang District, Zhanjiang Guangdong 524008, China
3. Graduate School of Kuroshio Science, Kochi University, 200 Otsu, Monobe, Nankoku, Kochi 738-8502, Japan