1. Divisions of Infectious Diseases, University of Michigan School of Medicine, Ann Arbor, MI, USA;
2. Department of Internal Medicine, University of Michigan School of Medicine, Ann Arbor, MI, USA;
3. Division of Infectious Diseases, Veterans Affairs Ann Arbor Healthcare System, Ann Arbor, MI, USA;
4. Department of Gastroenterology, University of Michigan School of Medicine, Ann Arbor, MI, USA;
5. Department of Microbiology and Immunology, University of Michigan School of Medicine, Ann Arbor, MI, USA