1. a Zabolotny Institute of Microbiology and Virology of National Academy of Sciences of Ukraine , Kyiv , Ukraine
2. b Institute of Geochemistry of the Environment of National Academy of Sciences of Ukraine , Kyiv , Ukraine
3. c Macaulay Institute, Craigiebuckler , Aberdeen , AB15 8QH , Scotland , UK
4. d Division of Molecular and Environmental Microbiology, College of Life Sciences , University of Dundee , Dundee , DD1 5EH , Scotland , UK