1. a Environmental Microbiology Group, Pacific Northwest National Laboratory , Richland , Washington , USA
2. b Department of Geosciences , Princeton University , Princeton , New Jersey , USA
3. c Geology Department , Carleton College, Northfield , Minnesota , 55057 , USA
4. d Department of Biology , New Mexico Institute of Mining and Technology , Socorro , New Mexico , USA
5. e Department of Biomedical Sciences Biology , The Florida State University , Tallahassee , Florida , 32306 , USA