1. a Consejo Superior de, Investigaciones Cientificas , Instituto de Recursos Naturales y Agrobiologia , Apdo. 257, Salamanca , 37071 , Spain
2. b Department Plant Ecology and Evolutionary Biology , University of Utrecht , P.O. Box 800, 84, TB Utrecht , 3508 , The Netherlands
3. c Section GVK, AB‐DLO , Research Institute for Agrobiology and Soil Fertility , Bornsesteg, 65, Postbus 14, Wageningen , The Netherlands