1. Botany Department, Faculty of Science, Tanta University, Tanta, Egypt
2. Department of Biology, College of Science and Arts at Khulis, University of Jeddah, Jeddah, Saudi Arabia
3. Genetic Engineering and Biotechnology Research Institute, University of Sadat City, Sadat City, Egypt
4. Botany Department, Faculty of Science, Menoufia University, Shibin El Kom, Egypt