1. Centre for Ecology, Evolution and Evironmental Changes (CE3C), Faculty of Sciences, University of Lisbon, 1749-016 Lisbon, Portugal
2. Conservation Science, Royal Botanic Gardens, Kew, Richmond, Surrey, TW9 3DS, UK
3. School of Biological Sciences, University of Reading, Whiteknights, Reading, Berkshire RG6 6BX, UK
4. School of Plant Biology, University of Western Australia, Crawley, WA 6009, Australia
5. Comparative Plant and Fungal Biology, Royal Botanic Gardens, Kew, Richmond, Surrey, TW3 3DS, UK