1. a Department of Geosciences, Geobiology Group , University of Göttingen , Goldschmidtstr. 3, 37077, Göttingen, Germany E-mail:
2. b Department of Fisheries and Marine Biology , University of Bergen , P.O. Box 7800, Høyteknologisenteret, 5020, Bergen, Norway E-mail:
3. c Institute of Biogeochemistry and Marine Chemistry , University of Hamburg , Bundesstr. 55, 20146, Hamburg, Germany E-mail:
4. d Research Centre Ocean Margins , University of Bremen , P.O. Box 330440, 28334, Bremen, Germany E-mail: