1. Academy of Agriculture and Forestry Sciences of Qinghai University (Qinhai Academy of Agriculture and Forestry Sciences), Xining, China
2. State Key Laboratory of Plateau Ecology and Agriculture, Qinghai University, Xining, China
3. Institute of Plant Virology, Fujian Agriculture and Forestry University, Fuzhou, China
4. Key Laboratory of the Tibet Plateau Biotechnology, Ministry of Education, Xining, China