1. Key Laboratory of Tropical Marine Bio-resources and Ecology, Guangdong Key Laboratory of Marine Materia Medica, Innovation Academy of South China Sea Ecology and Environmental Engineering, South China Sea Institute of Oceanology, Chinese Academy of Sciences, Guangzhou, China
2. University of the Chinese Academy of Sciences, Beijing, China
3. Southern Marine Science and Engineering Guangdong Laboratory (Guangzhou), Nansha District, Guangzhou, China