1. Biology Department, Faculty of Sciences, Chouaïb Doukkali University, El Jadida, Morocco
2. Laboratory of Microbial Biotechnologies, Agrosciences and Environment, Faculty of Sciences Semlalia, Cadi Ayyad University, Marrakech, Morocco
3. Department of Botany and Microbiology, College of Science, King Saud University, Riyadh, Saudi Arabia
4. Department of Agricultural and Environmental Sciences, Milan State University, Milan, Italy