1. Department of Biology, College of Sciences, Northern Border University, Arar Saudi Arabia
2. Laboratory of Plant Biotechnology, National Institute of Applied Sciences and Technology, Univesrty of Carthage, Tunis Cedex, Tunisia
3. Faculty of Science, Department of Biology, King Abdulaziz University, Jeddah, Saudi Arabia
4. Faculty of Science, Botany Department, South Valley University, Qena, Egypt