1. Department of Biological Sciences, College of Sciences, Northern Border University, Arar, Saudi Arabia
2. Laboratory of Plant Biotechnology, Department of Biology, National Institute of Applied Science and Technology, University of Carthage, Tunis, Tunisia
3. Laboratory of Natural Substances LR10INRAP02, National Institute of Research and Physico-Chemical Analyses, Biotechpole of Sidi Thabet, Ariana, Tunisia