1. Key Laboratory of Plant Resources Conservation and Sustainable Utilization/Guangdong Provincial Key Laboratory of Digital Botanical Garden, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou China
2. University of Chinese Academy of Sciences, Beijing, China
3. School of Pharmacy, Guangxi University of Chinese Medicine, Nanning, China
4. Key Laboratory of Vegetation Restoration and Management of Degradation Ecosystems/CAS Engineering Laboratory for Vegetation Ecosystem Restoration on Islands and Coastal Zones, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou, China