1. Tropical Crops Genetic Resources Institute, Chinese Academy of Tropical Agricultural Sciences/Tropical Wild Plant Gene Resource, Ministry of Agriculture/Key Laboratory of Biology and Cultivation of Herb Medicine (Haikou), Ministry of Agriculture and Rural Affairs, Haikou, P.R. China
2. Hainan Provincial Engineering Research Center for Tropical Medicinal Plants, Haikou, P.R. China
3. Key Laboratoory of South Subtropical Plant Diversity, Fairy Lake Botanical Garden, Shenzhen and Chinese Academy of Sciences, Shenzhen, P.R. China