1. Key Laboratory of Southern Subtropical Plant Diversity, Shenzhen Fairy Lake Botanical Garden, Chinese Academy of Sciences, 160 Xianhu Road, Luohu District, Shenzhen 518004, P.R. China
2. Key Laboratory of Plant Resources Conservation and Sustainable Utilization, South China Botanical Garden, Chinese Academy of Sciences, 723 Xingke Road, Tianhe District, Guangzhou 510650, P.R. China
3. The Key Laboratory of Chemical Biology, Guangdong Province, Division of Life Science, Graduate School at Shenzhen, Tsinghua University, Shenzhen 518055, P.R. China
4. Key Laboratory of Vegetation Restoration and Management of Degraded Ecosystems Ecosystems, South China Botanical Garden, Chinese Academy of Sciences 510650, P.R. China