1. Department of Biodiversity Conservation, Southwest Forestry University, Kunming, China;
2. Department of Life Science, Southwest Forestry University, Kunming, China;
3. National Plateau Wetlands Research Center, Southwest Forestry University, Kunming, China;
4. Department of Gardening and Horticulture, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Mengla, China