1. College of Biological Resource and Food Engineering, Qujing Normal University, Qujing, China
2. Key Laboratory of Yunnan Province Universities of the Diversity and Ecological Adaptive Evolution for Animals and Plants on YunGui Plateau, Qujing Normal University, Qujing, China
3. School of Resources and Environment, Baoshan University, Yunnan, China
4. Center for Integrative Conservation, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Yunnan, China