1. Jiangsu Key Laboratory of Marine Bioresources and Environment, Huaihai Institute of Technology, Lianyungang, China;
2. Jiangsu Key Laboratory of Marine Biotechnology, Huaihai Institute of Technology, Lianyungang, China;
3. Co-Innovation Center of Jiangsu Marine Bio-industry Technology, Lianyungang, China;
4. Simon F. S. Li Marine Science Laboratory School of Life Sciences, The Chinese University of Hong Kong, Shatin, Hong Kong, China